Its about a young woman,Andrea Sachs(Anne Hathaway) who wants the job of a journalist but ends up getting the job as Miranda Priestly's (Meryl Strepp)assistant. I think that was a BIG mistake because Miranda is a lady who has temper issues and Andrea had to do what ever humiliating task Miranda gave her.But due to the lack in time that Andrea gets to spend with her boyfriend, Nate(Adrian Grenier) he breaks up with her on the night that Miranda was taking Adrea to Paris with her for "Fall Fashion Week" and there she meets Christian Thompson(Simon Baker)who openly flirts with her.
At Adrea's friend Lily's (Tracie Thoms) art gallery exhibit Lily spots Adrea kissing Christian.In the end Adrea ditches her job and takes a job for a journalist while she gets back together with Nate . Nate gets a job as a chef in Bosten. Miranda smiles in the end that Adrea quit her job.